Every person living in any state or area requires a certain amount of infrastructure to meet their needs. More people, with more infrastructure equates to less habitat that is available for wildlife. With less habitat and more infrastructure there will generally be less wildlife available per individual living in the more populated states.
You constantly refer to the high harvest rates in 2b and 5C, yet now you claim that infra structure reduces the carrying capacity when the exact opposite is true You claim the habitat in 2G is controlling the herd with only 12 PS DPSM , but in developed areas like 2B it takes a harvest of 12 DPSM just to keep the herd stable.