Some good ideas.

Tried swapping one Y harness ends on the axles - no difference - that should have made it lean the other way, no?

Took off one of the cams - they both lean.

Rolled the axle and it appears straight. Could put an indicator on it, but with this amount of lean, it should be noticeable.

No slop in the cam bearing. Some minor slop in the four limb holes.