It might be that your paper is wrong, or perhaps you are. According to the DCNR website, they completed the southern Sproul flyovers and have the results posted.
http://www.dcnr.state.pa.us/forestry/deer/deersurvey_D10.aspx

What they found was not 16 deer per square mile, but a bit over 8. If you want facts, look at the page. If not, no problem with me.

Also, for those interested, here is the website for the Army Corps of Engineers study of the deer densities on Federal land at lake Raystown.
http://raystown.nab.usace.army.mil/LandManagement/deer_management.htm

"and the truth shall set you free."